The shannon weiner diversity index is a common way of showing that diversity involves not only numbers of different species, but also how well each of these species is represented in different habitats. Shannon weiner index the shannon index is affected by both the number of species and their equitability, or evenness. Bis 2b lab 6 graphs number of species vs panel age 8 7 6. Dpsi log pi, where s is the total number of species in the sample, i is the total number of individuals in. Margalef index, evenness index and shannon wiener index.
The shannonweaver diversity index is based on communication theory. A biological community has an attribute which we can call. Measuring diversity has been of historical significance and due to the obvious declines in habitat diversity this data is still valuable. Lets compute the shannon weiner diversity index for the same hypothetical community in the previous example. Shannon wiener index is defined and given by the following function. The shannon weiner index is most sensitive to the number of species in a sample, so it is usually considered to be biased toward measuring species richness.
The methods are brie y described, and the equations used them are given often in more detail than in their help pages. For our uses, this order could be characterized by the number of species andor the number of individuals in each species, within our sample plot. On some occasions it is called the shannon wiener index and on other occasions it is called the shannon weaver index. It turns out that the mathematical relationships hold true whether one is dealing with molecules in solution or species in an ecological community. View enhanced pdf access article on wiley online library html view download pdf for offline viewing. Shannon weiner index of diversity there are many methods that ecologists use to calculate species diversity. Quantitative measures of diversity, site similarity. Demonstration of how to calculate the shannonwiener diversity index in microsoft excel. We will then consider one measure used to compare diversity among communities, jaccard. Learn vocabulary, terms, and more with flashcards, games, and other study tools. If i obtain the shannonweiner diversity index as 2. The template works under windows os and excel 2010 xlsx extension. This study clearly shows that the comparison of diversity index is needed to set up the sample plots at the same size.
The intuitive significance of this index can be described as follows. Lets compute the shannonweiner diversity index for the same hypothetical community in the previous example. First, enter the number of species, and then enter the name you wish to give the species, if available, and the given populations for each of the species. Diversity indices like the shannon entropy shannon wiener index and the ginisimpson index are not themselves diversities. Sample values s 60,10,25,1,4 number of species n 5 first, let us calculate the sum of the given values. The actual formula for the shannon weiner diversity index is. We suggest an explanation for this dual use of terms and in so doing we offer a tribute to the late claude shannon who passed away on 24 february 2001.
Maximum species evenness was recorded for cyanophyceae and minimum for bacillariophyceae. Application of shannons index to study diversity with. The diversity calculator is an excel template that allows you to calculate alpha, beta and gamma diversity for a set samples input data, and to analyze similarities between the samples based on partitioning diversity in alpha and beta diversity. The shannon index is a measurement used to compare diversity between habitat samples. Diversity indices that combine both richness and abundance i.
View lab report bis 2b lab 6 graphs from bis 2b at university of california, davis. Pi in pi where pi s n s number of individuals of one species n total number of all individuals in the sample in logarithm to base e measurement of species richness margalefs index was used as a simple measure of. Pages 334338, in tested studies for laboratory teaching, volume. In fact, in the late 1940s shannon had built on the work of wiener. Maximum and minimum species diversity shannon weiner h were found 2. Shannon index is a commonly used diversity index that takes into account. The shannon wiener diversity index is one measure that we will use to try to draw information from samples in the field.
Biodiversity calculator this calculator is free to use and is designed for biologists, ecologists, teachers, and students needing to quickly calculate the biodiversity indexes of an ecosystem. Like simpsons index, shannon s index accounts for both abundance and evenness of the species present. Anyway, if we needed to create a network of long term ecological studies it. The most common measures of biodiversity are species richness, simpsons index and shannon wiener index. A greater number of species and a more even distribution both increase diversity as measured by h. Simpson diversity and the shannonwiener index as special. When all types in the dataset of interest are equally common, all pi values equal 1 r, and the shannon index hence takes the value ln r. In particular, for a random sample, we can use shannon s index of diversity aka as shannon weiner s index, which is defined as where n i is the number of observations from the sample in the i th of k nonempty categories and n is the sample size. Shannon s diversity index shannon s diversity index is simply the ecologists name for the communication entropy introduced by claude shannon. Dpsi log pi, where s is the total number of species in the sample, i is the total. This tutorial explains how to calculate the shannon wiener diversity index and evenness.
A tribute to claude shannon 19162001 and a plea for. Example diversity based on shrub dominance of a clay loam and a sandy loam site in south texas. A value for evenness approaching zero reflects large differences in abundance of species, whereas an evenness of one means all species are equally abundant. Statistics shannon wiener diversity index tutorialspoint. An example of the shannon wiener diversity index for science students. Both of these transformations serve to make the index increase as diversity intuitively increases, and although both are used, d 2 is more common magurran 2004. Historically, the index has been used to measure the effects of habitat quality such as effects of pollution effluents. The shannon weiner value h can range from no diversity at 0. The efficiency of indices of richness, evenness and. Species richness, diversity and abundance of some decapod crustaceans in coastal waters of ondo state, south west, nigeria. Sample values s 60,10,25,1,4 number of species n 5.
The shannonweiner species diversity index find, read and cite all the. In this system species are different only when their abundance. First, let us calculate the sum of the given values. The proportion of species i relative to the total number of species p i is calculated, and then multiplied by the natural logarithm of this. Species richness, diversity and abundance of some decapod. Calculate the shannon diversity index and evenness for these sample values. Interestingly shannon, a physicist, developed the index as a formula for measuring the entropy of matter in the universe.
The diversity index was calculated by using the shannon wiener diversity index 1949. Shannon index the idea behind this index is that the diversity of a community is similar to the amount of information in a code or message. The efficiency of indices of richness, evenness and biodiversity in the investigation of species diversity. Biodiversity calculator for the simpson and shannon indexes. This makes the shannon index and the simpson diversity or gini coefficient special cases of a more general index. Pdf on jan 1, 2006, kathleen a nolan and others published beachcomber biology. The shannon diversity index h is another index that is commonly used to characterize species diversity in a community. Number of species vs panel age 8 7 6 5 number of species 4 3 2 1 0 2 3 4 5 6 7 8. According to local occurrence status, there were 12 species ranked as very abundant, 19 abundant, 62 very common, 16 common and 41 fairly common. The shannonweiner species diversity index association for.
D 2 is closely related to d 1, being the inverse of simpsons original index simpson 1949. There were, however, 7 species, in each category, recorded as uncommon. This diversity measure is based on information theory. Simpson diversity and the shannon wiener index as special cases of a generalized entropy. The shannon wiener index is defined as is the natural logarithm of the total number of species. Recent interest in conservation biology has generated a strong focus on how to measure biodiversity in plant and animal communities. A measure used by ecologists when a system contains too many individuals for each to be identified and examined. Introduction, simpsons index and shannonweiner index. This comparison can be between two different habitats or a comparison of one habitat over time. All you do is count of the number of species found in a community e. H sumpi lnpi ehh max where, sum summation pi numbe of individuals of species itotal number of samples s number of species or species richness h max maximum diversity possible e evenesshh max. In the shannon index, p is the proportion nn of individuals of one particular species found n divided by the total number of individuals found n, ln is the natural log. This is by far the most widely used diversity index.
Shannon wiener index of diversity information index a measure used by ecologists when a system contains too many individuals for each to be identified and examined. The simpson index is a dominance index because it gives more weight to common or dominant species. Shannon wiener index of biodiversity and the simpson index of evenness showed contradictory results in the study area 199697 and 200809 figure 3 figure 4. It is different from species richness in that unlike richness it also shows community composition and takes into account the relative abundance of species that are present in the community. Shannon wiener index is the most widely index which used in marine biodiversity. Which site has the greatest diversity as estimated by the shannon weiner index h. Pages 334 338, in tested studies for laboratory teaching, volume. Values of the shannon diversity index for real communities typically fall between 1.
Key wordsdiversity indices, shannon index, shannon wiener index, shannon weaver index, species richness, species diversity. The maximum diversity hmax of a sample is found when all species are equally abundant. The uncertainty is measured by the shannon function h this term is the measure corresponding to the entropy concept defined by. The shannonwiener diversity index is one measure that we will use to try to draw information from samples in the field. A tribute to claude shannon 19162001 and a plea for more rigorous use of species richness, species diversity and the shannon wiener index. This is the simplest of all the measures of species diversity. A diversity index is the measure of species diversity in a given community.
Species diversity concepts describing communities species. Shannons diversity index real statistics using excel. They are just indices of diversity, in the same way that the diameter of a sphere is an index of its volume but is not itself the volume. The more unequal the abundances of the types, the larger the weighted geometric mean of the pi values.
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